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What does "toe the line" mean?

Also, toe the mark. Meet a standard, abide by the rules, as in The new director will make us toe the line, I'm sure, or At daycare Brian has to toe the mark, but at home his mother's quite lenient. This idiom refers to runners in a race placing their toes on the starting line and not moving until the starting signal. Its figurative use dates from the early 1800s.
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